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There are currently 15 recognised subspecies of great tit: [10]. The great tit is large for a tit at The nominate race P. The breast is bright lemon-yellow and there is a broad black mid-line stripe running from the bib to vent.

There is a dull white spot on the neck turning to greenish yellow on the upper nape. The rest of the nape and back are green tinged with olive.

The wing-coverts are green, the rest of the wing is bluish-grey with a white wing-bar. The tail is bluish grey with white outer tips.

The plumage of the female is similar to that of the male except that the colours are overall duller; the bib is less intensely black, [10] as is the line running down the belly, which is also narrower and sometimes broken.

There is some variation in the subspecies. The plumage of P. It is also slightly smaller, with a smaller bill but longer tail.

The situation is similar for the two related subspecies in the Turkestan tit group. The colour of the male bird's breast has been shown to correlate with stronger sperm, and is one way that the male demonstrates his reproductive superiority to females.

Higher levels of carotenoid increase the intensity of the yellow of the breast its colour, and also enable the sperm to better withstand the onslaught of free radicals.

The great tit is, like other tits, a vocal bird , and has up to 40 types of calls and songs. The calls are generally the same between the sexes, but the male is much more vocal and the female rarely calls.

Soft single notes such as "pit", "spick", or "chit" are used as contact calls. A loud "tink" is used by adult males as an alarm or in territorial disputes.

One of the most familiar is a "teacher, teacher", often likened to a squeaky wheelbarrow wheel, which is used in proclaiming ownership of a territory.

One explanation for the great tit's wide repertoire is the Beau Geste hypothesis. The eponymous hero of the novel propped dead soldiers against the battlements to give the impression that his fort was better defended than was really the case.

Similarly, the multiplicity of calls gives the impression that the tit's territory is more densely occupied than it actually is.

Whether the theory is correct or not, those birds with large vocabularies are socially dominant and breed more successfully.

The great tit has a wide distribution across much of Eurasia. It can be found across all of Europe except for Iceland and northern Scandinavia , including numerous Mediterranean islands.

The great tit occupies a range of habitats. It is most commonly found in open deciduous woodland , mixed forests , forest edges and gardens.

In dense forests, including conifer forests it prefers forest clearings. In northern Siberia it lives in boreal taiga. In North Africa it rather resides in oak forests as well as stands of Atlas cedar and even palm groves.

In the east of its range in Siberia, Mongolia and China it favours riverine willow and birch forest. Riverine woodlands of willows, poplars are among the habitats of the Turkestan subspecies, as well as low scrubland , oases ; at higher altitudes it occupies habitats ranging from dense deciduous and coniferous forests to open areas with scattered trees.

The great tit is generally not migratory. Pairs will usually remain near or in their territory year round, even in the northern parts of their range.

Young birds will disperse from their parents' territory, but usually not far. Populations may become irruptive in poor or harsh winters, meaning that groups of up to a thousand birds may unpredictably move from northern Europe to the Baltic and also to Netherlands, Britain, even as far as the southern Balkans.

The great tit was unsuccessfully introduced into the United States; birds were set free near Cincinnati, Ohio between and but failed to become established.

Suggestions that they were an excellent control measure for codling moths nearly led to their introduction to some new areas particularly in the United States of America, however this plan was not implemented.

Great tits are primarily insectivorous in the summer, feeding on insects and spiders which they capture by foliage gleaning.

Seeds and fruit usually come from deciduous trees and shrubs, like for instance the seeds of beech and hazel. Where it is available they will readily take table scraps, peanuts and sunflower seeds from bird tables.

Large food items, such as large seeds or prey, are dealt with by "hold-hammering", where the item is held with one or both feet and then struck with the bill until it is ready to eat.

Using this method, a great tit can get into a hazelnut in about twenty minutes. When feeding young, adults will hammer off the heads off large insects to make them easier to consume, and remove the gut from caterpillars so that the tannins in the gut will not retard the chick's growth.

Great tits combine dietary versatility with a considerable amount of intelligence and the ability to solve problems with insight learning, that is to solve a problem through insight rather than trial and error.

This is the first time a songbird has been recorded preying on bats. The tits only do this during winter when the bats are hibernating and other food is scarce.

Great tits are monogamous breeders and establish breeding territories. Females are likely to disperse to new territories if their nest is predated the previous year.

If the pair divorces for some reason then the birds will disperse, with females travelling further than males to establish new territories.

Great tits are seasonal breeders. The exact timing of breeding varies by a number of factors, most importantly location.

Most breeding occurs between January and September; in Europe the breeding season usually begins after March. In Israel there are exceptional records of breeding during the months of October to December.

The amount of sunlight and daytime temperatures will also affect breeding timing. Great tits are cavity nesters, breeding in a hole that is usually inside a tree, although occasionally in a wall or rock face, and they will readily take to nest boxes.

The nest inside the cavity is built by the female, and is made of plant fibres, grasses, moss, hair, wool and feathers.

The number in the clutch is often very large, as many as 18, but five to twelve is more common. Clutch size is smaller when birds start laying later, and is also lower when the density of competitors is higher.

Insularity also affects clutch size, with great tits on offshore islands laying smaller clutches with larger eggs than mainland birds.

The female undertakes all incubation duties, and is fed by the male during incubation. The timing of hatching, which is best synchronised with peak availability of prey, can be manipulated when environmental conditions change after the laying of the first egg by delaying the beginning of incubation, laying more eggs or pausing during incubation.

The chicks, like those of all tits , are hatched unfeathered and blind. Once feathers begin to erupt, the nestlings are unusual for altricial birds in having plumage coloured with carotenoids similar to their parents in most species it is dun-coloured to avoid predation.

The nape is yellow and attracts the attention of the parents by its ultraviolet reflectance. This may be to make them easier to find in low light, or be a signal of fitness to win the parents' attention.

This patch turns white after the first moult at age two months, and diminishes in size as the bird grows. Feeding of the fledgeling may continue after independence, lasting up to 25 days in chicks from the first brood, but as long as 50 days in the second brood.

Inbreeding depression occurs when the offspring produced as a result of a mating between close relatives show reduced fitness.

The reduced fitness is generally considered to be a consequence of the increased expression of deleterious recessive alleles in these offspring.

In natural populations of P. The Eurasian sparrowhawk is a predator of great tits, with the young from second broods being at higher risk partly because of the hawk's greater need for food for its own developing young.

It was originally a specialist tit flea, but the dry, crowded conditions of chicken runs enabled it to flourish with its new host.

Although these beetles often remain in deserted nests, they can only breed in the elevated temperatures produced by brooding birds, tits being the preferred hosts.

Incidences of fatal competition are more frequent when nesting times overlap, and climate change has led to greater synchrony of nesting between the two species and flycatcher deaths.

Having killed the flycatchers, the great tits may consume their brains. The great tit is a popular garden bird due to its acrobatic performances when feeding on nuts or seed.

Its willingness to move into nest boxes has made it a valuable study subject in ornithology ; it has been particularly useful as a model for the study of the evolution of various life-history traits, particularly clutch size.

The great tit has generally adjusted to human modifications of the environment. It is more common and has better breeding success in areas with undisturbed forest cover, but it has adapted to human modified habitats.

It can be very common in urban areas. In areas with low frequency background noise pollution, the song has a higher frequency than in quieter areas.

While there have been some localised declines in population in areas with poorer quality habitats, its large range and high numbers mean that the great tit is not considered to be threatened, and it is classed as least concern on the IUCN Red List.

From Wikipedia, the free encyclopedia. Linnaeus , Parus major. Biology Letters. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis.

Tomus I. Editio decima, reformata in Latin.

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The plumage of P. It is also slightly smaller, with a smaller bill but longer tail. The situation is similar for the two related subspecies in the Turkestan tit group.

The colour of the male bird's breast has been shown to correlate with stronger sperm, and is one way that the male demonstrates his reproductive superiority to females.

Higher levels of carotenoid increase the intensity of the yellow of the breast its colour, and also enable the sperm to better withstand the onslaught of free radicals.

The great tit is, like other tits, a vocal bird , and has up to 40 types of calls and songs. The calls are generally the same between the sexes, but the male is much more vocal and the female rarely calls.

Soft single notes such as "pit", "spick", or "chit" are used as contact calls. A loud "tink" is used by adult males as an alarm or in territorial disputes.

One of the most familiar is a "teacher, teacher", often likened to a squeaky wheelbarrow wheel, which is used in proclaiming ownership of a territory.

One explanation for the great tit's wide repertoire is the Beau Geste hypothesis. The eponymous hero of the novel propped dead soldiers against the battlements to give the impression that his fort was better defended than was really the case.

Similarly, the multiplicity of calls gives the impression that the tit's territory is more densely occupied than it actually is. Whether the theory is correct or not, those birds with large vocabularies are socially dominant and breed more successfully.

The great tit has a wide distribution across much of Eurasia. It can be found across all of Europe except for Iceland and northern Scandinavia , including numerous Mediterranean islands.

The great tit occupies a range of habitats. It is most commonly found in open deciduous woodland , mixed forests , forest edges and gardens. In dense forests, including conifer forests it prefers forest clearings.

In northern Siberia it lives in boreal taiga. In North Africa it rather resides in oak forests as well as stands of Atlas cedar and even palm groves.

In the east of its range in Siberia, Mongolia and China it favours riverine willow and birch forest. Riverine woodlands of willows, poplars are among the habitats of the Turkestan subspecies, as well as low scrubland , oases ; at higher altitudes it occupies habitats ranging from dense deciduous and coniferous forests to open areas with scattered trees.

The great tit is generally not migratory. Pairs will usually remain near or in their territory year round, even in the northern parts of their range.

Young birds will disperse from their parents' territory, but usually not far. Populations may become irruptive in poor or harsh winters, meaning that groups of up to a thousand birds may unpredictably move from northern Europe to the Baltic and also to Netherlands, Britain, even as far as the southern Balkans.

The great tit was unsuccessfully introduced into the United States; birds were set free near Cincinnati, Ohio between and but failed to become established.

Suggestions that they were an excellent control measure for codling moths nearly led to their introduction to some new areas particularly in the United States of America, however this plan was not implemented.

Great tits are primarily insectivorous in the summer, feeding on insects and spiders which they capture by foliage gleaning. Seeds and fruit usually come from deciduous trees and shrubs, like for instance the seeds of beech and hazel.

Where it is available they will readily take table scraps, peanuts and sunflower seeds from bird tables. Large food items, such as large seeds or prey, are dealt with by "hold-hammering", where the item is held with one or both feet and then struck with the bill until it is ready to eat.

Using this method, a great tit can get into a hazelnut in about twenty minutes. When feeding young, adults will hammer off the heads off large insects to make them easier to consume, and remove the gut from caterpillars so that the tannins in the gut will not retard the chick's growth.

Great tits combine dietary versatility with a considerable amount of intelligence and the ability to solve problems with insight learning, that is to solve a problem through insight rather than trial and error.

This is the first time a songbird has been recorded preying on bats. The tits only do this during winter when the bats are hibernating and other food is scarce.

Great tits are monogamous breeders and establish breeding territories. Females are likely to disperse to new territories if their nest is predated the previous year.

If the pair divorces for some reason then the birds will disperse, with females travelling further than males to establish new territories.

Great tits are seasonal breeders. The exact timing of breeding varies by a number of factors, most importantly location.

Most breeding occurs between January and September; in Europe the breeding season usually begins after March. In Israel there are exceptional records of breeding during the months of October to December.

The amount of sunlight and daytime temperatures will also affect breeding timing. Great tits are cavity nesters, breeding in a hole that is usually inside a tree, although occasionally in a wall or rock face, and they will readily take to nest boxes.

The nest inside the cavity is built by the female, and is made of plant fibres, grasses, moss, hair, wool and feathers.

The number in the clutch is often very large, as many as 18, but five to twelve is more common. Clutch size is smaller when birds start laying later, and is also lower when the density of competitors is higher.

Insularity also affects clutch size, with great tits on offshore islands laying smaller clutches with larger eggs than mainland birds.

The female undertakes all incubation duties, and is fed by the male during incubation. The timing of hatching, which is best synchronised with peak availability of prey, can be manipulated when environmental conditions change after the laying of the first egg by delaying the beginning of incubation, laying more eggs or pausing during incubation.

The chicks, like those of all tits , are hatched unfeathered and blind. Once feathers begin to erupt, the nestlings are unusual for altricial birds in having plumage coloured with carotenoids similar to their parents in most species it is dun-coloured to avoid predation.

The nape is yellow and attracts the attention of the parents by its ultraviolet reflectance. This may be to make them easier to find in low light, or be a signal of fitness to win the parents' attention.

This patch turns white after the first moult at age two months, and diminishes in size as the bird grows.

Feeding of the fledgeling may continue after independence, lasting up to 25 days in chicks from the first brood, but as long as 50 days in the second brood.

Inbreeding depression occurs when the offspring produced as a result of a mating between close relatives show reduced fitness.

The reduced fitness is generally considered to be a consequence of the increased expression of deleterious recessive alleles in these offspring.

In natural populations of P. The Eurasian sparrowhawk is a predator of great tits, with the young from second broods being at higher risk partly because of the hawk's greater need for food for its own developing young.

It was originally a specialist tit flea, but the dry, crowded conditions of chicken runs enabled it to flourish with its new host. Although these beetles often remain in deserted nests, they can only breed in the elevated temperatures produced by brooding birds, tits being the preferred hosts.

Incidences of fatal competition are more frequent when nesting times overlap, and climate change has led to greater synchrony of nesting between the two species and flycatcher deaths.

Having killed the flycatchers, the great tits may consume their brains. The great tit is a popular garden bird due to its acrobatic performances when feeding on nuts or seed.

Its willingness to move into nest boxes has made it a valuable study subject in ornithology ; it has been particularly useful as a model for the study of the evolution of various life-history traits, particularly clutch size.

The great tit has generally adjusted to human modifications of the environment. It is more common and has better breeding success in areas with undisturbed forest cover, but it has adapted to human modified habitats.

It can be very common in urban areas. In areas with low frequency background noise pollution, the song has a higher frequency than in quieter areas.

While there have been some localised declines in population in areas with poorer quality habitats, its large range and high numbers mean that the great tit is not considered to be threatened, and it is classed as least concern on the IUCN Red List.

From Wikipedia, the free encyclopedia. Linnaeus , Parus major. Biology Letters. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis.

Tomus I. Editio decima, reformata in Latin. Holmiae Laurentii Salvii. Cassell's Latin Dictionary 5th ed. London: Cassell Ltd. The ornithology of Francis Willughby of Middleton in the county of Warwick, esq.

London, United Kingdom: A. Proceedings of the Royal Society B. RA, ed. Check-list of birds of the world. Volume Cambridge, Massachusetts: Museum of Comparative Zoology.

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